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The chief activator of AMPK is the serine-threonine tumor suppressor kinase LKB1, which contributes to phosphorylation of AMPK to activate energy sensors (235, 238)

Mandel SA, Amit T, Weinreb O, Reznichenko L, Youdim MB

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This can lead to improved energy levels and a reduction in the frequency and severity of fatigue-related symptoms

This phosphorylation, which occurs via the JNK pathway, is considered to be crucial as it modifies FOXO4, preparing it for its role in the cell's nucleus