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PDI consists of four domains ( a, b, b, a ), two of which ( a and a ) contain thioredoxin-related reactive CXXC motifs that facilitate disulfide exchange ( b and b domains are not catalytically active, but act as chaperones for substrate binding (Klappa et al., 1998)

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GHK and its copper complexes penetrate the stratum corneum and can be delivered transdermally in creams, gels or skin patches [ 49 , 50 ]